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Calicioids of Temperate Regions
0.01 (development)
Eric B. Peterson
 

Chaenothecopsis


Chaenothecopsis may be the largest genus among the Calicioids and potentially undescribed members are surprisingly common.  The challenge is that they are so small and with limited morphological diversity that species can be difficult to distinguish.  The majority appear only as small black stalked ascomata with no thallus.  I examine a sample first, marking (usually with insect pins) any variants in form that may suggest multiple species within the sample, then start plucking ascomata to look at under a compound scope. Creating a single key for all species of Chaenothecopsis is rather cumbersome, both for the key writer and for key users.  I have broken it down by some basic groups in the following table, followed by separate keys for each group.

Taxa covered (79):

  Chaenothecopsis "cascades"
  Chaenothecopsis "ghostly"
  Chaenothecopsis "greenfoot"
  Chaenothecopsis "hobit"
  Chaenothecopsis "imperatrice"
  Chaenothecopsis "klamath"
  Chaenothecopsis "purple"
  Chaenothecopsis aeruginosa
  Chaenothecopsis amurensis
  Chaenothecopsis arthoniae
  Chaenothecopsis asperopoda
  Chaenothecopsis australis
  Chaenothecopsis brevipes
  Chaenothecopsis caespitosa
  Chaenothecopsis caucasica
  Chaenothecopsis cinerea
  Chaenothecopsis claydenii
  Chaenothecopsis consociata
  Chaenothecopsis debilis
  Chaenothecopsis diabolica
  Chaenothecopsis dolichocephala
  Chaenothecopsis edbergii
  Chaenothecopsis epithallina
  Chaenothecopsis Eugenia
  Chaenothecopsis exilis
  Chaenothecopsis exserta
  Chaenothecopsis fennica
  Chaenothecopsis formosa
  Chaenothecopsis golubkovae
  Chaenothecopsis haemtopus
  Chaenothecopsis heterospora
  Chaenothecopsis himalayensis
  Chaenothecopsis hospitans
  Chaenothecopsis hyrcana
  Chaenothecopsis irregularis
  Chaenothecopsis jaczevskii
  Chaenothecopsis kalbii
  Chaenothecopsis khayensis
  Chaenothecopsis kilimanjaroensis
  Chaenothecopsis koerberi
  Chaenothecopsis lecanactidis
  Chaenothecopsis leifiana
  Chaenothecopsis mediarossica
  Chaenothecopsis montana
  Chaenothecopsis nana
  Chaenothecopsis nigra
  Chaenothecopsis nigropedata
  Chaenothecopsis nivea
  Chaenothecopsis norstictica
  Chaenothecopsis ochroleuca
  Chaenothecopsis oregana
  Chaenothecopsis parasitaster
  Chaenothecopsis pilosa
  Chaenothecopsis pusilla
  Chaenothecopsis pusiola
  Chaenothecopsis quintralis
  Chaenothecopsis rappii
  Chaenothecopsis resinicola
  Chaenothecopsis retinens
  Chaenothecopsis rubescens
  Chaenothecopsis rubina
  Chaenothecopsis sagenidii
  Chaenothecopsis sanguinea
  Chaenothecopsis savonica
  Chaenothecopsis shefflerae
  Chaenothecopsis sinensis
  Chaenothecopsis sitchensis
  Chaenothecopsis subparoica
  Chaenothecopsis tasmanica
  Chaenothecopsis thujae
  Chaenothecopsis tibellii
  Chaenothecopsis transbaikalica
  Chaenothecopsis trassii
  Chaenothecopsis tristis
  Chaenothecopsis ussuriensis
  Chaenothecopsis vainioana
  Chaenothecopsis vinosa
  Chaenothecopsis viridialba
  Chaenothecopsis viridireagens

Warning: keys are incomplete.

 
Key Substrate soaked with Resin Spore Septation
A yes any
B no yes
C no no

Validation script believes the following taxa are missing from the keys:

  1. Chaenothecopsis "klamath"
  2. Chaenothecopsis caucasica
  3. Chaenothecopsis exserta
  4. Chaenothecopsis formosa
  5. Chaenothecopsis heterospora
  6. Chaenothecopsis hyrcana
  7. Chaenothecopsis leifiana
  8. Chaenothecopsis mediarossica
  9. Chaenothecopsis rappii
  10. Chaenothecopsis sinensis
  11. Chaenothecopsis transbaikalica

A. Key to Resinicolous Chaenothecopsis:


  1. 1z. Ascomata with a golden to greenish pruina.
    1. 2a. Spores septate, 7.6 - 10.1 μm in length. Known from various conifers in North America, in both Pacific and Atlantic regions.Chaenothecopsis edbergii
    2. 2b. Spores non-septate.
      1. 3a. Known only from Sequoia and Sequoiadendron (and apparently endemic to California).Mycocalicium sequoiae
      2. 3b. Spores 3.5 - 5.3 μm; known only from Pinus; pruina may vary from greenish to white.Chaenothecopsis resinicola
  2. 1z. [step].[leaf]
    1. 4a. Spores septate.
      1. 5a. Capitulum with a thick white pruina; spores being extruded through the pruina to form black dots over the surface. Known only from Abies in North America where it occurs primarily in branch scars.Chaenothecopsis nigripunctata
      2. 5b. Capitulum not as above (epruinose or thinly pruinose).
        1. 6c. White to greenish pruina.Chaenothecopsis asperopoda
    2. 4b. Spores non-septate (at least the majority of mature spores).
      1. 7a. Spore mass distinctly brown in color.Bruceomyces castoris
      2. 7b. Spore mass distinctly black, even if covered by a pruina.
        1. 8a. K-.
          1. 9a. On viscin of Mistletoe (Tristerix); known only from high altitudes in Ecuador and Peru.Mycocalicium viscinicola
          2. 9b. On other substrates.
            1. 10a. Spores minute, 3.5 - 5.5 μm long; stalk usually with a rough surface and a greenish to whitish pruina.Chaenothecopsis resinicola
            2. 10b. Spores larger.
              1. 11a. On hardwoods, known from tropical Africa.Chaenothecopsis khayensis
              2. 11b. On conifers, known from the Pacific Northwest of North America.
                1. 12a. Growing on distinct resin; ascomata typically thick-stalked; spores 6.7 - 7.5 μm long with broadly rounded ends and a greenish color.Chaenothecopsis montana
                2. 12b. Growing on well exposed bark with just a hint of resin; ascomata typically slender; ascospores 6 - 8 μm long, allantoid (somewhat pointed ends), with a dark brown pigmentation. This species is typically not treated as resinicolous.Chaenothecopsis nana
        2. 8b. K (any color).
          1. 13a. K red to purple. (Also known as C. oregana.).Chaenothecopsis diabolica
          2. 13b. K green.
Step number 1 has 7 options!
Step number 6 has 6 options!


B. Key to Non-resinicolous Chaenothecopsis with Septate Spores:


  1. 1a. Ascomata reacting red, purple, or green to KOH (may fade quickly) as viewed in squash-mount with compound microscope.
    1. 2a. Color reaction to KOH is red or purple (may be fade quickly).
      1. 3a. Color reaction wine-red to deep purple.
        1. 4a. Color reaction typically purplish wine-red and fading quickly (you may even need to watch carefully as the KOH wave passes through the specimen). Ascomata often found on resin-impregnated wood.Chaenothecopsis pusiola
        2. 4b. Color reaction typically deep purple. Known only from wood of Quercus in southern Oregon, U.S.A.Chaenothecopsis "purple"
      2. 3b. Color reaction fire-engine red to orangish-red.
        1. 5a. [step].[leaf]
        2. 5b. [step].[leaf]
    2. 2b. Color reaction to KOH is green.
      1. 6a. Obligate parasite on Chaenotheca chrysocephala.Chaenothecopsis consociata
      2. 6b. Not an obligate parasite on Chaenotheca chrysocephala.
        1. 7a. Stalk in water mount appearing intensely red; spores narrowly ellipsoidal, 7 - 8 μm long.Chaenothecopsis rubina
        2. 7b. Upper part with +/- reddish tinge; epithecium and stalk mainly brown or olive-brown but stalk sometimes reddish in parts; spore _ long.Chaenothecopsis viridireagens
  2. 1b. Ascomata without a color reaction to KOH as viewed in squash-mount with compound microscope, or if a color reaction occurs, then turning dingy-yellow or brown.
    1. 8a. Saprophytic vs parasitic needs lookup.
    2. 8d. Growing parasitically on a lichen thallus (sometimes other calicioids). Note that it should be clear that it is growing from the thallus, not from wood but surrounded by a lichen thallus; sometimes leprose lichens will cover substrates where Chaenothecopsis is growing saprophytically.
    3. 8e. Growing saprophytically from wood or bark. Algal colonies may be present, or not, within the substrate.
      1. 11a. Ascomata with white or bluish-white dusty pruina on either capitulum or stalk. Spore septation strong.
        1. 12a. Ascomata very small, ≤ 0.5 mm tall; capitula typically spherical and entirely encrusted in pruina. Known from deep fissures and beetle tunnels in bark of Pseudotsuga menziesii.Chaenothecopsis "ghostly"
        2. 12b. Ascomata of variable size, often > 0.5 mm tall; capitula relatively tall and narrow; pruina often with a bluish tinge.
      2. 11b. Ascomata lacking a dusty pruina (though sometimes a pale waxy sheen may form on the excipulum). Strength of spore septation somewhat weak except for one species (C. debilis).
        1. 14a. Ascospores typically 7 - 9 μm. Stalk and capitulum often 1 mm or taller, and with greenish pigments.Chaenothecopsis aeruginosa
        2. 14b. Ascospores typically 6 - 7 μm long. Ascomata typically < 1 mm tall (often significantly smaller). Pigments variable.
          1. 15a. Ascomata small, ≤ 0.5 mm tall, darkly pigmented in squash mount (often deeply green but sometimes brown to black). Ascomata formed within the bark between flakes or other cracks.Chaenothecopsis "greenfoot"
          2. 15b. Ascomata smallish, but often > 0.5 mm tall; pigments visible in squash mount primarily brown or even slightly reddish. Ascomata growing where exposed to light.
            1. 16a. HNO3+ red. [Stalk not swelling in KOH? - I need to double check this.] A variable taxon in terms of pruina and ascomata density. I suspect that it actually represents two species: one epruinose, typically sparse ascomata on bark of Quercus or Pseudotsuga; the other pruinose, typically forming dense colonies over Quercus bark.Chaenothecopsis debilis
            2. 16b. HNO3-. Stalk swelling strongly in KOH. Pigmentation variable. Substrates variable. Sometimes on substrates with algae present. Likely a cluster of cryptic speciation.Chaenothecopsis pusilla
Step number 3 has 8 options!
Step number 8 has 5 options!
Step number 9 has 18 options!


 

C. Key to Non-resinicolous Chaenothecopsis with Non-septate Spores


  1. 1a. Ascomata reacting red, purple, or green to KOH (may fade quickly) as viewed in squash-mount with compound microscope.
    1. 2a. Color reaction to KOH is red or purple (may be fade quickly).
    2. 2b. Color reaction to KOH is green.
      1. 4a. Spores allantoid, with a strong brown pigment. Ascomata short-stalked, typically ≤ 0.3 mm tall. Excipulum typically constricted at the rim.Chaenothecopsis "imperatrice"
      2. 4b. Spores not allantoid.
  2. 1b. Ascomata without a color reaction to KOH as viewed in squash-mount with compound microscope, or if a color reaction occurs, then turning dingy-yellow or brown.
    1. 6a. Growing on dung.Chaenothecopsis quintralis
    2. 6b. Growing on other substrates.
      1. 7z. (presumed K- but not yet tested). Very small (< 0.3 mm), stalk pale to white, capitula black. Ascospores variable in size and pointiness.Chaenothecopsis "hobit"
Step number 3 has 5 options!
Step number 5 has 6 options!
Step number 7 has 13 options!


 

Chaenothecopsis "cascades"
ID=71
Type(s):
Synonyms:
Chaenothecopsis "cascades" ascomata. California, Trinity County.

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Chaenothecopsis "ghostly"
ID=233
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Chaenothecopsis "greenfoot"
ID=135
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Chaenothecopsis "hobit"
ID=251
Type(s):
Synonyms:
Chaenothecopsis "hobit" Photographed with a 10x objective, focal stacked from 94 images.
Chaenothecopsis "hobit" Ascospores photographed at 400x.  Note variability in length and pointiness.

Habit and Morphology:

Small with a slighly fuzzy stalk.


Anatomy:
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Hidden within bark of Pseudotsuga menziesii.


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Chaenothecopsis "imperatrice"
ID=236
Type(s):
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Unspecified collections:
Chaenothecopsis sp. nov. at ca. 9x. Chaenothecopsis sp. nov. from Imperatrice, near Ashland, OR. Scot Loring # SCL16935. Ca. 9x.
Chaenothecopsis sp. nov. "Imperatrice", Scot Loring #  SCL16935 Squash mount at 100x showing hyaline base.
Chaenothecopsis sp. nov Squash mount (pressed) with red pigmentation before addition of KOH. 100x. Scot Loring # SCL16935
Chaenothecopsis sp. nov. Squash mount (pressed) with red pigmentation reacting green with addition of KOH. 100x. Scot Loring # SCL16935
Chaenothecopsis sp. nov. Squash mount (pressed) after red pigmentation reacted green with addition of KOH. 100x. Scot Loring # SCL16935
Chaenothecopsis sp. nov. Ascus showing Chaenothecopsis-form of tip, with an apical canal (albeit not very clearly - better photo needed). 400x. Scot Loring # SCL16935
Chaenothecopsis sp. nov. Ascospore. 400x. Scot Loring # SCL16935

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Found by Scot Loring at "Imperatrice" near Ashland, OR, specimen number SCL16925.


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Chaenothecopsis "klamath"
ID=235
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Chaenothecopsis "purple"
ID=239
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Unspecified collections:
Chaenothecopsis "purple" Stacked macrophotography.
Chaenothecopsis "purple" Stacked macrophotography.
Chaenothecopsis "purple" Stacked macrophotography.
Chaenothecopsis "purple" K+ purple reaction (moderately strong example)
Chaenothecopsis "purple" K+ purple reaction (weak example)
Chaenothecopsis "purple" Ascospores photographed at 400x.

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Chaenothecopsis aeruginosa
ID=64
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Chaenothecopsis amurensis Titov
ID=65
Type(s): Titov & Tibell 1993
Synonyms:
Habit and Morphology: Associated with Trentepohlia. Thallus immersed. Ascomata small, 0.2-0.4 mm tall.
Anatomy: Ascomata epruinose. Stalk outwardly of periclinal hyphae; more pale and irregularly interwoven inward. Asci cylindrical to narrowly clavate; apex thick with a fine canal. Ascospores on-septate, ellipsoidal to allantoid, brown, 4.5 - 8 um X 1.8 - 3.9 um, smooth to very minutely ornamented.
Chemistry: K-, N-
Habitat: Moist deciduous or mixed Pinus - deciduous forests. Reported only from bark of Phellodendron amurense.
Biogeography: Russian far east.
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External Information: References: Titov & Tibell 1993.
Chaenothecopsis arthoniae Tibell
ID=66
Type(s): (Tibell 1998)
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Chaenothecopsis asperopoda Titov
ID=67
Type(s):

Titov & Tibell (1993)


Synonyms:
Habit and Morphology:

On conifer exudate (Picea) without algae.

Thallus not described.

Ascomatal stature medium to large, 0.5 - 2.5 mm tall, straight to flexuous; sometimes with a reddish color to the lower excipulum and upper stalk.


Anatomy:

Stalk with a rough texture and often covered by a greenish or whitish pruina. Capitulum black. Excipulum poorly developed.

[left off - NEED TO COMPLETE]


Chemistry:

K+R->G (reddish pigments intensifying, then becoming aeruginose); N-.


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External Information: References: Selva & Tibell 1999; Titov & Tibell 1993.
Chaenothecopsis australis Tibell
ID=68
Type(s): (Tibell 1998)
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Chaenothecopsis brevipes Tibell
ID=69
Type(s):

Tibell 1987


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External Information: References: Tibell 1987; Titov & Tibell 1993.
Chaenothecopsis caespitosa (Phillips) D. Hawksw.
ID=70
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Chaenothecopsis caucasica Titov
ID=72
Type(s):

Titov 2006


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External Information: References: Titov 2006.
Chaenothecopsis cinerea Tibell
ID=73
Type(s): (Tibell 1998)
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Chaenothecopsis claydenii Selva & Tuovila
ID=243
Type(s): Selva & Tuovila 2017
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Chaenothecopsis consociata (N?dv.) A.F.W. Schmidt
ID=74
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External Information: References: Titov & Tibell 1993.
Chaenothecopsis debilis (Turner & Borrer ex Sm.) Tibell
ID=75
Type(s):
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Unspecified collections:
Chaenothecopsis debilis (one form). Typical epruinose form found on Quercus garryana bark, often in just sparse, scattered patches and on small trees (but old - stunted growth). S. Loring # SCL16982, on Quercus garryana (resin soaked bark), North Bank HMA,  Chasm Creek.<
Chaenothecopsis debilis ascospores. S. Loring # SCL16982, on Quercus garryana (resin soaked bark), North Bank HMA,  Chasm Creek.
Chaenothecopsis debilis reaction to Nitric Acid. S. Loring # SCL16982, on Quercus garryana (resin soaked bark), North Bank HMA,  Chasm Creek.
Chaenothecopsis debilis ascomata in squash mount. California, Trinity County.
Chaenothecopsis debilis ascospores. California, Trinity County.

Habit and Morphology:

Saprophytic, described by Tibell as only on wood (but western North American material on Quercus bark is commonly aligned with this taxon), usually without algae.

Thallus immersed.

Ascomata medium sized, 0.4 - 1.2 mm tall, black, with or without a 'pruina' on the excipulum that sometimes extends down the stalk. The 'pruina' described by Tibell (1987) as "cottony hyphae" and rare but distinctive when present. In western North America the pruina seems almost exclusively restricted to colonies on Quercus bark which tend to be either very sparse, small, individual ascomata without pruina, or dense colonies of stout ascomata with abundant pruina. These patterns are suggestive of multiple poorly distinguished lineages.


Anatomy:

Stalk outwardly of pigmented periclinal hyphae, inwardly pale and hyphae more branched. Stalk wine red in section.

Asci cylindrical, tip penetrated with a fine canal.

Mature ascospores medium brown, distinctly septate, 6.5 - 9.8 um, smooth to minutely areolate.


Chemistry:

K- or K+ grayish read in stalk; N+ reddish pigments intensifying toward violet red (sometimes faint in western North America).


Habitat:
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Distribution is likely global, or nearly so, with literature reports from Australia, Eurasia, and North America at the least.


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Good description in Tibell 1987 (Australasian).


External Information: References: Tibell 1987; Titov & Tibell 1993.
Chaenothecopsis diabolica Rikkinen & Tuovila
ID=76
Type(s): Tuovila et al. 2011; still in my key as C. oregana
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Chaenothecopsis dolichocephala Titov
ID=77
Type(s): (Selva 2010)
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Chaenothecopsis edbergii Selva & Tibell
ID=78
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Chaenothecopsis edbergii ascomata and mycelial mat. Tillamook County, Oregon.

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External Information: References: Selva & Tibell 1999.
Chaenothecopsis epithallina Tibell
ID=79
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Unspecified collections:
Chaenotheca trichialis paracitized by Chaenothecopsis epithallina Riley 0766; from 35mm slide

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Chaenothecopsis Eugenia Titov
ID=80
Type(s): 2001
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Chaenothecopsis exilis Tibell
ID=81
Type(s):

(1999): (Selva & Tibell 1999)


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Unspecified collections:
Chaenothecopsis exilis. (I believe - but havent experienced this species much.) Collection J. Chambers #WINJC41.

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External Information: References: Selva & Tibell 1999.
Chaenothecopsis exserta (Nyl.) Tibell
ID=241
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Habit and Morphology: Parasitic on Haemotomma ochroleucum. Ascomata sessile, or nearly so.
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Chemistry: K+ red.
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Chaenothecopsis fennica (M. Laurila) Tibell
ID=82
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Chaenothecopsis formosa Titov (2006)
ID=83
Type(s):

Titov 2006


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External Information: References: Titov 2006.
Chaenothecopsis golubkovae Tibell & Titov
ID=84
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External Information: References: Titov & Tibell 1993.
Chaenothecopsis haemtopus Tibell
ID=85
Type(s):

1987


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External Information: References: Titov & Tibell 1993.
Chaenothecopsis heterospora Titov
ID=86
Type(s):

(2006): Titov 2006


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External Information: References: Titov 2006.
Chaenothecopsis himalayensis (R?s?nen) Tibell & Titov
ID=87
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Chaenothecopsis hospitans (Th. Fr.) Tibell
ID=88
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Chaenothecopsis hyrcana Titov
ID=89
Type(s):

(2006): Titov 2006


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External Information: References: Titov 2006.
Chaenothecopsis irregularis Titov
ID=90
Type(s):

-1991


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External Information: References: Titov & Tibell 1993.
Chaenothecopsis jaczevskii Titov
ID=91
Type(s):

Titov (2006) includes Protocalicium jaczevskii


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Chaenothecopsis kalbii Tibell & K. Ryman
ID=92
Type(s): (Lendemer et al. 2008c)
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Chaenothecopsis khayensis Rikkinen & Tuovila
ID=244
Type(s): Tuovila et al. 2011
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Chaenothecopsis kilimanjaroensis Temu and Tibell
ID=250
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External Information: References: Temu et al. 2019.
Chaenothecopsis koerberi (Nadv.) Tibell
ID=94
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Chaenothecopsis lecanactidis Tibell
ID=95
Type(s): (Tibell 1998)
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Chaenothecopsis leifiana Titov
ID=96
Type(s): -2004
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Chaenothecopsis mediarossica Titov & Gudovicheva
ID=99
Type(s):

(2006): Titov 2006


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Chaenothecopsis montana Rikkinen
ID=100
Type(s): (2003): Rikkinen 2003; Tuovila et al 2011
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Chaenothecopsis nana Tibell
ID=101
Type(s):

-1979


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Chaenothecopsis nigra Tibell
ID=102
Type(s): (1987) (Harris & Lendemer 2005, Spribille & Bj?rk 2008)
Synonyms:
Unspecified collections:
Chaenothecopsis nigra Hardman #6113, on loan through Daphne Stone. Ascomata, showing size, and form (first and fourth are most typical). Synthesis of several photos, with focus stacking and with scale maintained; photographed at ca. 10x.
Chaenothecopsis nigra Hardman #6113, on loan through Daphne Stone. Squash mount showing hyphal arrangement that is irregular to slightly periclinal; photographed at 100x.
Chaenothecopsis nigra Hardman #6113, on loan through Daphne Stone. Spores photographed at 400x.
Chaenothecopsis nigra (cf) California, Napa County.
Chaenothecopsis nigra (cf) ascospores. California, Napa County.

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Chaenothecopsis nigropedata Tibell
ID=103
Type(s): -1987
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Chaenothecopsis nivea (F. Wilson) Tibell
ID=104
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Chaenothecopsis norstictica R. C. Harris
ID=105
Type(s): 1995
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Chaenothecopsis ochroleuca (K?rber) Tibell & K. Ryman
ID=106
Type(s):

-1995


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Chaenothecopsis oregana Rikkinen
ID=107
Type(s): (2003) = C. diabolica and C. zebrina: Rikkinen 2003; Tuovila et al. 2011
Synonyms:
Unspecified collections:
Chaenothecopsis oregana. Collection E. Burnett #L3442.

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Chaenothecopsis parasitaster (Bagl. & Car.) D. Hawksw.
ID=108
Type(s): -1978
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Chaenothecopsis pilosa Tibell & Kalb
ID=109
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Chaenothecopsis pusilla (Ach.) A.F.W. Schmidt
ID=110
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Chaenothecopsis pusiola (Ach.) Vain
ID=111
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Chaenothecopsis quintralis Messuti, Amico, Lorenzo, & Vidal-Russel
ID=249
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Chaenothecopsis rappii (N?dv.) R. C. Harris
ID=112
Type(s):

may be in my key under Mycocalicium


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Chaenothecopsis resinicola Tibell & Titov
ID=113
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Chaenothecopsis retinens (Nyl.) Tibell
ID=114
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Chaenothecopsis rubescens Vainio
ID=116
Type(s):
Synonyms:
Chaenothecopsis rubescens ascomata in squash mount.
Sclerophora peronella ascomata. With Chaenothecopsis rubescens.
Chaenothecopsis rubescens ascospores. Darkly pigmented and non-septate, though I would not call them allantoid as the ends are well rounded.

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Chaenothecopsis rubina Tibell
ID=117
Type(s): 1982
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Chaenothecopsis sagenidii Tibell
ID=118
Type(s):

-1987


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Chaenothecopsis sanguinea Tibell
ID=119
Type(s): -1987
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Chaenothecopsis savonica (R?s?nen) Tibell
ID=120
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Chaenothecopsis shefflerae (Samuels & D.E. Buchanan) Tibell
ID=121
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Chaenothecopsis sinensis Titov
ID=122
Type(s):

(2006): Titov 2006


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Chaenothecopsis sitchensis Rikkinen
ID=123
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Chaenothecopsis subparoica (Nyl.) Tibell
ID=124
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Chaenothecopsis tasmanica Tibell
ID=125
Type(s):

(Tibell 1987


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Chaenothecopsis thujae Rikkinen
ID=248
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Chaenothecopsis tibellii Titov
ID=126
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Chaenothecopsis transbaikalica Titov
ID=127
Type(s):

(2006): Titov 2006


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Chaenothecopsis trassii Titiov
ID=128
Type(s):

(1998): Titov 2006


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Chaenothecopsis tristis (Koerb.) Titov
ID=129
Type(s): (1998): Titov & Tibell 1998 listed in Tibell?s checklist (?99), Europe
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Chaenothecopsis ussuriensis Titov
ID=131
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Chaenothecopsis vainioana
ID=132
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Chaenothecopsis vinosa Titov
ID=133
Type(s): (2001): Titov 2001
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Chaenothecopsis viridialba (Kremp.) A.F.W. Schmidt
ID=134
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Chaenothecopsis viridireagens (N?dv.) A.F.W. Schmidt
ID=136
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Count of taxa = 79

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